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By contrast, DPP-4 inhibitors constrain the DPP-4 enzyme that deactivates a variety of bioactive peptides including GLP-1 and glucose-dependent insulinotropic polypeptide, enhancing their endogenous levels and thereby increasing glucose-dependent insulin secretion and decreasing glucagon secretion, like GLP-1RAs [Citation5]

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Accordingly, in INS-1 832/3 and MIN6B1 cells, accumulation of cAMP (with a low concentration of 3-isobutyl-1-methylxanthine (IBMX) to prevent cAMP degradation) was greater with exendin-phe1 than with exendin-4 and exendin-asp3 (Fig