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Acknowledgments We thank Haiyan Yu, Xiaomin Zhao and Yuyu Guo from Core Facilities for Life and Environmental Sciences of SKLMT (State Key Laboratory of Microbial Technology, Shandong University) for the assistance in LSM900 Super Resolution Laser Scanning Confocal Microscope analysis

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Received: Mar 18, 2025

Primary murine PDAC cells were derived from Kras tm1Tyj : Kras LSL-G12D/+ + Trp53 tm1Brn : Trp53 fl/fl + Ptf1a tm1(Cre)Hnak : Ptf1a Cre-ex1/+ (KPC) compound mutant mouse tumors and were prepared as fragments for outgrowth culture