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The promoter of the hCG- gene contains tandem cyclic adenosine monophosphate (cAMP) response elements (CREs), an upstream regulatory sequence, and a junctional response element (JRE), which specify trophoblast expression of the subunit gene.16, 17, 18, 19 In trophoblast cells, the CREs bind CRE-binding proteins such as AP-1, the recruitment of which can be stimulated by epidermal growth factor and adenylate cyclase, and JREs bind placenta-specific trans-acting factors, such as the homeobox factor Dix 3.11, 16, 17, 18, 19 Together, this array of regulatory elements, in conjunction with tissue-specific expression of trans-acting factors, endows the promoter of the hCG- gene with trophoblast specificity and maximal activity.17 The hCG- genes evolved from the luteinizing hormone (LH) subunit and have acquired new regulatory elements that favor their expression in the placenta rather than the pituitary gland

doi: 10.1515/znc-2024-0116 3 ShangSYangHQuLFanDDengJ

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